Norway's Handball Federation (NHF) said it would pay the fines but would "continue to fight to change the rules for clothing, so that players can play in the clothes they are comfortable with".

Norway's sports minister Abid Raja said attitudes needed to change, calling the situation "completely ridiculous".

In September, government ministers from Denmark, Finland, Iceland, Norway and Sweden had asked the IHF to review its uniform rules "in accordance with gender equality".

Meanwhile, Norway-based Australian activist Talitha Stone had been gathering signatures for a petition that called for the uniform rules to be scrapped.

Over the weekend, Ms Stone told her supporters her campaign had been successful, after 61,000 signed the petition.

In IHF's latest beach handball rule book, dated 3 October, there was no mention of bikinis.

The updated rules say female beach handball players can now wear "body fit" tank tops and " short tight pants", as opposed to crop tops and bikini bottoms.

Male athletes are allowed to wear shorts that are "not too baggy" but must remain 10cm (3.9in) above the kneecap.

The BBC has requested comment from the IHF.

Ms Stone said she hoped the rule change would be the "beginning of the end of sexism and objectification of women and girls in sport".

She said all women and girls should "be free to participate in sport without fear of wardrobe malfunctions and sexual harassment". (BBC)